Fred Keeley was born and raised in Winnipeg where he completed an undergraduate degree in Chemistry and a PhD in Pharmacology at the University of Manitoba. He spent two years as a Post-Doctoral fellow at the British Agricultural Research Council research establishment near Bristol, England, followed by a Post- Doctoral fellowship in the Department of Biochemistry at the Hospital for Sick Children. He joined the staff of the Research Institute of the Hospital for Sick Children in 1973, and is currently a Senior Scientist in the Molecular Structure and Function Program. From 1995 to 2013 he served as Associate Chief of Research at the Hospital for Sick Children, and held the Heart and Stroke Foundation of Ontario/Robert M. Freedom Chair in Cardiovascular Science at the Hospital for Sick Children from 2003 to 2008 and from 2008 to 2013. Dr. Keeley was first cross-appointed to the University of Toronto in 1975, and is currently Professor in the Department of Biochemistry and the Department of Laboratory Medicine and Pathobiology.