My research has addressed a wide variety of topics related to the behavior, ecology, evolution, and conservation of Pacific salmon, trout, and char, and their ecosystems. Some of my long-standing interests are the patterns of migration and homing by salmon, and the mechanisms that underlie these behavior patterns, and also the evolutionary aspects of homing – the formation of locally adapted populations, and gene flow arising from straying. This work has involved natural populations, populations transplanted to other locations, and the recolonization of habitat after dam removal or modification. I also have a long-standing interest in predator-prey interactions and especially the ecological and evolutionary aspects of predation by bears on salmon, as well as the ecological and evolutionary effects of human activities on salmon and trout, such as selective fishing and artificial propagation in hatcheries.